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Despite the theft of four prototype Gundams by ZAFT, the Earth Alliance continues its research and constructs new units featuring advanced technologies. One such unit is the GAT-X370 Raider Gundam, which is designed as a transforming mobile suit similar to the GAT-X303 Aegis Gundam. One of the new technologies implemented in the Raider Gundam is Trans-Phase armor, which is an improved version of Phase Shift armor. Whereas Phase Shift armor utilizes a 'power up' mode denoted by color and a 'power down' mode denoted by gray, Trans-Phase armor is designed to only activate at the point of impact and thus preserve energy. This energy can then be diverted to power-consuming energy weapons. Also, to keep up with the abilities of Coordinators, the Alliance designates pilot Clotho Buer as a 'biological CPU' rather than a pilot and requires him to take drugs to be able to effectively use his high performance mobile suit. Unfortunately, these drugs have severe side effects after wearing off.

Like the Aegis Gundam, the Raider Gundam is equipped with weapons that are operable in its variable configurations. In mobile suit mode, the Raider Gundam features a twin machine gun mounted on its right arm. It also features a "Zorn" energy cannon in place of a mouth, which is operable in either configuration. For close combat, the Raider Gundam is also equipped with a "Mjolnir" hammer. While in flight mode, the Raider Gundam can utilize the "Ahura Mazda" cannons located on the claws, as well as several machine guns mounted on the nose and shoulders. The armaments featured in the Raider Gundam are a variation on the original design, the GAT-333 Raider Full Spec. The Raider Gundam is first deployed in combat during the Earth Alliance's attack on the Orb Union. While on Earth, the Raider Gundam also serves as a subflight unit for the ground unit GAT-X131 Calamity Gundam. Following the defeat of Orb, the Raider Gundam is taken to space and transferred to the Archangel class battleship Dominion under the supervision of Blue Cosmos leader Muruta Azrael. The Raider Gundam is destroyed near the war's end on September 26, C.E. 71, by Yzak Joule's GAT-X102 Duel Gundam.